Caudammina ovuloides (Grzybowski, 1901)



Fig. 42. Lectotype of Reophax ovuloides, from Grzybowski (1901)


ORIGINAL DESIGNATION: Reophax ovuloides Grzybowski, 1901

TYPE REFERENCE: Grzybowski, J., 1901, Otwornice warstw inoceramowych okolicy Gorlic. Rozprawy Wydziału Matematyczno-Przyrodniczego, Akademia Umiejętności w Krakowie, serya 2, vol. 41, p. 223, pl. 8, fig. 3. See also: Kaminski, M.A. & Geroch, S., 1993. A revision of foraminiferal species in the Grzybowski Collection. Grzybowski Foundation Special Publication, 1, 239-323, pl. 15. fig. 5a,b [Lectotype].

TYPE SPECIMEN: Not originally designated. A lectotype was disignated by Kaminski & Geroch (1993; pl. 15, fig. 5). This specimen was illustrated by Grzybowski (1901; pl. 8, fig. 3), and is preserved in the micropaleontological collections of the Jagiellonian University, Kraków, (no. UJ-133-P 2/30).

TYPE LEVEL: The lectotype specimen is from Paleocene variegated shales overlying the Inoceramian beds, Magura tectonic unit of the Polish Carpathians.

TYPE LOCALITY: The lectotype is from the Rogoyski well (30 m depth) at Michalówka, in the village of Siary, near Gorlice Poland.

DIAGNOSTIC FEATURES: Test free, pseudomultilocular, comprised of two or more rounded, pyriform or elongated pseudochambers without true internal partitions, connected by relatively thick, but short, tubular stolons. Wall thick, imperforate, finely agglutinated, usually well-silicified. Aperture at the open end of the tube at one or both ends of the test.

SIZE: The lectotype specimen is 1.09 mm in length. Single chambers may be up to 0.90 mm in length.

SUSPECTED SYNONYMS: None verified.

OBSERVED OCCURRENCES: Caudammina ovuloides was originally reported by Grzybowski (1901) as Reophax ovuloides from the Paleocene of the Polish Carpathians. Subsequent authors have either reported it as Hormosina ovuloides or combined it with Caudammina ovulum. It was recorded from the Paleogene of the Polish Carpathians (Jednorowska, 1975). Hanzlíková (1969, 1972) reported it from the Maastrichtian Frydek Formation and the Maastrichtian to Paleocene Istebna beds in Moravia; and Samuel (1977) recorded it from the Upper Cretaceous to Paleocene Inoceramian, Cisna, and Proc beds of the Carpathians in eastern Slovakia. De Klasz & De Klasz (1990) illustrated a specimen from the Danian of the Buntmergelserie in Bavaria. Beckmann (1994) reported it as Hormosinella ovulum from the mid Paleocene of Trinidad.
Caudammina ovuloides also occurs in Upper Cretaceous to Paleogene sediments of the North Atlantic region. It has been reported from the Maastrichtian at DSDP Site 543 by Hemleben & Troester (1984), from the Maastrichtian of ODP Site 641 (Moullade et al., 1988); from the Paleocene at ODP Site 900 (Kuhnt & Collins, 1996), from the Lower Campanian of the Hacho de Montejaque section in southern Spain (Kuhnt, 1990); and from the Coniacian-Santonian (Anguille Stage) to lowermost Maastrichtian (Point-Claire Stage) of offshore Gabon by Volat et al. (1996). We have observed it in the Campanian to Paleocene of Trinidad, in the ?Turonian through Campanian of offshore Norway, in the Paleocene of Zumaya Spain, in the Upper Paleocene of DSDP Site 112 in the Labrador Sea, in the Paleocene at Site 959 in the equatorial Atlantic, in the Paleocene Kamalapuram Formation of the Cauvery Basin, India, and in the Early to Middle Eocene "Glomospira assemblage" at ODP Site 643, Norwegian-Greenland Sea.

KNOWN STRATIGRAPHIC RANGE: ?Turonian - early Middle Eocene. Not observed above the Paleocene in the Atlantic region south of the Greenland-Scotland Ridge.

BATHYMETRY: Bathyal to abyssal.

REMARKS: Caudammina ovuloides differs from Caudammina ovula in possessing more pyriform chambers which taper towards the tube, and in having a connecting tube that is thicker, more robust, and shorter in proportion to the diameter of the test. In this respect, it is closer in morphology to the Recent species Reophanus oviculus (=Hormosina ovicula Brady, 1879). The length of the connecting tube is variable. Multichambered specimens of C. ovuloides are occasionally found, whereas in C. ovula the connecting stolon invariably breaks. Specimens of C. ovuloides from Zumaya Spain have a slightly coarser wall than specimens from other studied localities.
The lectotype of C. ovuloides displays a second chamber which is smaller than the first, although forms with larger second chambers are more typical. The smaller second chamber in this case may be the result of adverse growth conditions (the individual appears to have closed up the aperture at the wide basal portion of the test and moved into a smaller second chamber), or reproduction.

ILLUSTRATIONS: Plate 42 - Caudammina ovuloides (Grzybowski)
Fig. 1a-b. Paleocene, Siary Poland, Magura Unit of the Polish Carpathians, Lectotype; Fig. 2. Paleocene, Szklary Poland, Skole Unit of the Polish Carpathians; Fig. 3. Campanian, Węglówka Poland, Subsilesian Unit of the Polish Carpathians; Fig. 4. Paleocene, DSDP Site 112, Labrador Sea, Sample 112-16R-1, 85-87 cm; Fig. 5. Paleocene, Zumaya Spain; Fig. 6. Lower Paleocene, "Scaglia Rossa" facies, beach outcrop in Sopelana Spain; Fig. 7. Lower Paleocene, Lizard Springs Formation of Trinidad, well G-163, 4566'.